Comparison Between Genetic and Environmental Influences on Lumber Bending Properties in Young White Spruce
Keywords:
Modulus of elasticity, modulus of rupture, <i>Picea glauca</i>, studs, wood density, knotsAbstract
This study investigated variation in lumber bending properties of white spruce (Picea glauca [Moench] Voss) and its correlation with tree growth, wood density, and knot size and number. A total of 242 sample trees from 39 open-pollinated families harvested from 36-year-old provenance-progeny trials at two sites in Quebec, Canada through a thinning operation were processed. The results indicate that mechanical properties of lumber from young white spruce plantation-grown trees are low. It appears that low wood density, the occurrence of numerous knots, and a high proportion of juvenile wood are the main factors contributing to the low lumber stiffness and strength properties. The narrow-sense heritability for lumber stiffness was low to moderate, whereas that of strength was hardly different from zero. Thus environmental growing conditions highly influence white spruce wood mechanical properties. The results also revealed a strong negative correlation between stem volume and lumber stiffness and strength at the family means, which suggests that selection for volume would have an indirect negative effect on lumber quality. However, the absence of such significant correlation at the phenotypic level also suggests that mass selection with vegetative propagation would be a promising avenue for improving white spruce wood properties without having to give up gains in volume.References
American Society for Testing Materials (ASTM) D4761-02a/D198-02e1. 2002a. Standard test methods for mechanical properties of lumber and wood-base structural material/Standard test methods of static tests of lumber in structural sizes. ASTM International, West Conshohocken, PA.nAmerican Society for Testing Materials (ASTM) D1990-00. 2002b. Standard practice for establishing allowable properties for visually-graded dimension lumber from in-grade tests of full-size specimens. ASTM International, West Conshohocken, PA.nAmerican Society for Testing Materials (ASTM) 2915-03. 2003. Standard practice for evaluating allowable properties for grades of structural lumber. ASTM International, West Conshohocken, PA.nBao, F. C., Z. H. Jiang, X. M. Jiang, X. X. Lu, X. Q. Luo, and S. Y. Zhang. 2001. Differences in wood properties between juvenile wood and mature wood in 10 species grown in China. Wood Sci. Technol.35(4):363-375.nBarrett, J. D., and R. M. Kellogg. 1991. Bending strength and stiffness of second-growth Douglas-fir dimension lumber. Forest Prod. J.41(10):35-43.nBarrett, J. D., and W. Lau. 1994. Canadian lumber properties. Canadian Wood Council, Ottawa, Canada. 346 pp.nBeaulieu, J., B. Girard, and Y. Fortin. 2003. Study of geographical variation in kiln-drying behavior of plantation-grown white spruce. Wood Fiber Sci.35(1):56-67.nBecker, W. A. 1985. Manual of quantitative genetics. 4th edition. Academic Enterprises, Pullman, WA. 194 pp.nBendtsen, B. A. 1978. Properties of wood from improved and intensively managed trees. Forest Prod. J.28(10):61-72.nBiblis, E. J. 1990. Properties and grade yield of lumber from a 27-year-old slash pine plantation. Forest Prod. J.40(3): 21-24.nBiblis, E. J., H. Carino, R. Brinker, and C. W. McKee. 1995. Effect of stand density on flexural properties of lumber from two 35-year-old loblolly pine plantations. Wood Fiber Sci.27(1):25-33.nBiblis, E. J., H. Carino, R. Brinker, and C. W. McKee. 1997. Flexural properties of lumber from two 40-year-old loblolly pine plantations with different stand densities. Wood Fiber Sci.29(4): 375-380.nBier, H. 1985. Bending properties of structural timber from a 28-year-old stand of New Zealand Pinus radiata.N. Z. J. For. Sci.15(2):233-250.nBier, H. 1986. Log quality and the strength and stiffness of structural timber. N. Z. J. For. Sci.16(2):176-186.nBolghari, H. A., and V. Bertrand. 1984. Tables préliminaires de production des principales essences résineuses plantées dans la partie centrale du sud du Québec. Gouv. Quebec, Min. Energ. Ressour., Serv. Rech., Mem. Rech. For. No 79. 392 pp.nBurdon, R. D., R. A. J. Britton, and G. B. Walford. 2001. Wood stiffness and bending strength in relation to density in four native provenances of Pinus radiata.N. Z. J. For. Sci.31(1):130-146.nCanadian Council of Forest Ministers (CCFM). 2003. National Forestry Database Program http://nfdp.ccfm.org'>http://nfdp.ccfm.orgnCanadian Wood Council (CWC). 2001. Wood Design Manual 2001. Ottawa, ON. 900 pp.nCave, I. D., and J. C. F. Walker. 1994. Stiffness of wood in fast-grown plantation softwoods: The influence of microfibril angle. Forest Prod. J.44(5):43-48.nChui, Y. H. 1995. Grade yields and wood properties of Norway spruce (Picea abies (L.) Karst.) from the Maritimes. For. Chron.71(4):473-478.nCorriveau, A., J. Beaulieu, and F. Mothe. 1987. Wood density of natural white spruce populations in Quebec. Can. J. For. Res.17(7):675-682.nCorriveau, A., J. Beaulieu, F. Mothe., J. Poliquin, and J. Doucet. 1990. Densité et largeur de cernes des populations d'Épinettes blanches de la région forestière des Grands Lacs et du Saint-Laurent. Can. J. For. Res.20(2):121-129.nCorriveau, A., J. Beaulieu, and G. Daoust. 1991. Heritability and genetic correlations of wood characters of Upper Ottawa Valley white spruce populations grown in Quebec. For. Chron.67(6):698-705.nCown, D. J., J. Hebert, and R. Ball. 1999. Modelling Pinus radiata lumber characteristics. Part 1: Mechanical properties of small clears. N. Z. J. For. Sci.29(2):203-213.nDanborg, F. 1996. Juvenile wood in Norway and Sitka spruce: Anatomy, density, drying properties, visual grading and strength properties. Forskningsserien no. 18. Danish Forest and Landscape Research Institute, Horsholm, Denmark.nDhir, N. K. 1976. Stand, family, and site effects in Upper Ottawa Valley white spruce. Pages 88-97 in D. W. Einspahr, ed. Twelfth Lake States Forest Tree Improvement Conference. August 28-22, 1975. Petawawa, ON. USDA Forest Service, General Technical Report NC-26. St. Paul, MNnDieters, M. J., T. L. White, R. C. Littell, and G. R. Hodge. 1995. Application of approximate variances of variance components and their ratios in genetic tests. Theor. Appl. Genet.91:15-24.nFalconer, D. S. 1981. Introduction to quantitative genetics. Longman, London, UK, 340 pp.nForintek Canada Corp. 1995. Characteristics of Alberta commercial tree species. Properties of white spruce (Picea glauca (Moench) Voss). Alberta Department of Economic Development and Tourism. 16 pp.nGrant, D. J., A. Anton, and P. Lind. 1984. Bending strength, stiffness, and stress-grade of structural Pinus radiata: effect of knots and timber density. N. Z. J. For. Sci.14(3):331-348.nHarris, J. M., R. N. James, and M. J. Collins. 1976. Case for improving wood density in radiata pine. N. Z. J. For. Sci.5(3):347-354.nIvkovich, M., G. Namkoong, and M. Koshy. 2002. Genetic variation in wood properties of interior spruce. I. Growth, latewood percentage, and wood density. Can. J. For. Res.32(12):2116-2127.nJayawickrama, K. J. S., and M. J. Carson. 2000. A breeding strategy for the New Zealand radiata pine breeding cooperative. Silvae Genet.49(2):82-90.nJessome, A. P. 1977. Strength and related properties of woods grown in Canada. Forestry Tech. Rep. 21. Eastern Forest Products Laboratory. Ottawa, ON.nJozsa, L. A., and G. R. Middleton. 1994. A discussion of wood quality attributes and their practical implications. Forintek Canada Corp. Special Publ. No. SP-34. 42 pp.nKeith, C. T., and R. M. Kellogg 1981. The structure of wood. Chapter 3, Pages 41-70 in E. J. Mullins and T. S. McKnight, eds. Canadian woods. Their properties and uses. University of Toronto Press, Toronto, ON.nKennedy, R. W. 1995. Coniferous wood quality in the future: Concerns and strategies. Wood Sci. Technol.29: 321-338.nKiss, G., and F. C. Yeh. 1988. Heritability estimates for height for young interior spruce in British Columbia. Can. J. For. Res.18:158-162.nKliger I. R. 2000. Mechanical properties of timber products required by end-users. Proc. World Conference on Timber Engineering, July 31-August 3, 2000, Whistler, BC. 9 pp.nKretschmann, D. E., and B. A. Bendtsen. 1992. Ultimate tensile stress and modulus of elasticity of fast-grown plantation loblolly pine lumber. Wood Fiber Sci.24(2): 189-203.nLi, P., J. Beaulieu, A. Corriveau, and J. Bousquet. 1993. Genetic variation in juvenile growth and phenology in a white spruce provenance-progeny test. Silvae Genet.42(1):52-60.nLynch, M., and B. Walsh. 1998. Genetics and analysis of quantitative traits. Sinauer Associates Inc., Sunderland, MA. 980 pp.nMacPeak, M. D., L. F. Burkart, and D. Weldon. 1990. Comparison of grade, yield, and mechanical properties of lumber produced from young fast-grown and older slow-grown planted slash pine. Forest Prod. J.40(1):11-14.nMatheson, A. C., J. L. Yang, and D. L. Spencer. 1997a. Breeding radiata pine for improvement of sawn product value. Pages IV-19-IV-26 in S. Y. Zhang, R. Gosselin, and G. Chauret, eds. Timber management toward wood quality and end-product value. Proc. CTIA/IUFRO International Wood Quality Workshop, August 18-22, 1997, Quebec City, Canada.nMatheson, A. C., D. J. Spencer, J. G. Nyakuengama, J. Yang, and R. Evans. 1997b. Breeding for wood properties in radiata pine. Pages 169-179 in R. D. Burdon, and J. M. Moore, eds. IUFRO 97' Genetics of Radiata Pine. Proc. NZFRI-IUFRO Conference. 1-5 December 1997, Rotorura, New Zealand. New Zealand Forest Research Institute Bulletin No. 203.nMattsson, S. 2002. Effect of site preparation on stem growth and clear wood properties in boreal Pinus sylvestris and Pinus contorta. Acta Universitatis Agriculturae Sueciae Silvestria, 240. 139 p.nMcAlister, R. H., and H. R. Powers. 1994. A comparison of some physical and mechanical properties of full-sib and half-sib loblolly pines. Forest Prod. J.44(2):42-44.nMcKeand, S., T. Mullin, T. Byram, and T. White. 2003. Deployment of genetically improved loblolly and slash pines in the South. J. For.101(3):32-37.nMegraw, R., D. Bremer, G. Leaf, and J. Roers. 1999. Stiffness in loblolly pine as a function of ring position and height, and its relationship to microfibril angle and specific gravity. Pages 341-349 in G. Nepveu, ed. Workshop IUFRO S5.01-04. La Londe-Les-Maures, September 5-12, 1999. INRA, Nancy, France.nMilliken, G. A., and D. E. Johnson. 1984. The analysis of messy data, Vol. 1. Designed experiments. Van Nostrand Reinhold Co., New York, NY.nMoulton, R. J. and G. Hernandez. 2000. Tree planting in the United States - 1998. Tree Planters' Notes49(2): 23-26.nOkuma, M., N. Nakamura, and K. Yamada. 1984. Manufacture and performance of radiata pine plywood. II. Pruning effect on the properties of plywood. J. Jap. Wood Res. Soc.30(6):440-447.nOlson, R. A., and N. V. Poletika. 1947. Strength properties of plantation-grown coniferous woods. Conn. Agric. Exp. Stn. Bull. 511.nPearson, R. G., and R. C. Gilmore. 1971. Characterization of the strength of juvenile wood of loblolly pine (Pinus taeda L.). Forest Prod. J.21(1):23-30.nPearson, R. G., and R. C. Gilmore. 1980. Effect of fast growth rate on mechanical properties of loblolly pine. Forest Prod. J.30(5):47-54.nPorter, A. W. 1981. Strength and physical properties of wood. Chapter 4, Pages 71-96 in E. J. Mullins and T. S. McKnight, eds. Canadian woods. Their properties and uses. University of Toronto Press, Toronto, ON.nRochester, G. H. 1938. The strength of eastern Canadian spruce timbers in sizes shipped to the United Kingdom. Canada Forest Service. Circular 54.nSAS Institute Inc. 2001. SAS OnlineDoc®, version 8.02. SAS Institute Inc., Cary, NC.nSorensson, C. T., D. J. Cown, B. G. Ridoutt, and X. Tian. 1997. The significance of wood quality in tree breeding: a case study of radiata pine in New Zealand. Pages IV-35-IV-44 in S. Y. Zhang, R. Gosselin, and G. Chauret, eds. Timber Management Toward Wood Quality and End-Product Value. Proc. CTIA/IUFRO International Wood Quality Workshop, August 18-22, 1997, Quebec City, Canada.nStanturf, J. A., R. C. Kellison, F. S. Broerman, and S. B. Jones. 2003. Productivity of southern pine plantations. Where are we and how did we get here? J. For.101(3): 26-31.nXie, C.-Y., and A. D. Yanchuk. 2002. Genetic parameters of height and diameter of interior spruce in British Columbia. Forest Genet.9(1):1-10.nYanchuk, A. D., and G. K. Kiss. 1993. Genetic variation in growth and wood specific gravity and its utility in the improvement of interior spruce in British Columbia. Silvae Genet.42(2/3):141-148.nZhang, S. Y., G. Chauret, H. Q. Ren, and R. Desjardins. 2002. Impact of initial spacing on plantation Black spruce lumber grade yield, bending properties, and MSR yield. Wood Fiber Sci.34(3):460-475.nZhou, H., and I. Smith. 1991. Factors influencing bending properties of white spruce lumber. Wood Fiber Sci.23(4):483-500.nZobel, B. J., and J. B. Jett. 1995. Genetics of wood production. Springer-Verlag GmbH & Co. KG, Berlin, Germany.nZobel, B. J., and J. R. Sprague. 1998. Juvenile wood in forest trees. Springer Series in Wood Science. T. E. Timell, ed. Springer-Verlag, New York, NY.nZobel, B. J., and J. P. van Buijtenen. 1989. Wood variation. Its causes and control. Springer-Verlag GmbH & Co. KG, Berlin, Germany.n
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