Intraclonal Variation in Wood Density of Trembling Aspen in Alberta
Keywords:Wood density, <i>Populus tremuloides</i> Michx, intraclonal variation, specific gravity
AbstractFour trees from each of three putative clones of trembling aspen (Populus tremuloides Michx.) at one site in north-central Alberta were sampled to determine the patterns of wood density variation within stems and within clones. Sample disks were removed at five heights from each tree to examine variation among cardinal directions and across the southern radius at each height. Although only three clones were sampled, there were significant differences (0.05 level) among clones. Wood density tends to be high at the bottom of the tree, decreases to a minimum at midheight, then increases again near the top of the tree. In the radial direction, wood density is high near the pith (at all heights), decreases, then increases again in the mature wood zone (after rings 15-20+). Average wood density values within the twelve stems varied from 0.348 g/cc to 0.402 g/cc.
Barnes, B. V. 1966. The clonal growth habit of American aspens. Ecol.47(3):439-447.nBarnes, B. V. 1969. Natural variation and delineation of clones of Populus tremuloides and P. grandidentata in northern lower Michigan. Silv. Genet.18(4):130-141.nBrown, I. R. 1961. Clonal variation in specific gravity in Populus tremuloides. Unpublished M.Sc. thesis, State College of Forestry at Syracuse, New York.nBrown, I. R., and F. A. Valentine. 1963. Natural variation in specific gravity and fiber length in Populus tremuloides clones. 10th. Northeastern For. Tree Imp. Conf. Proc. Pp. 25-41.nEinspahr, D. W. 1972. Wood and fiber production from short rotation stands. In Aspen; symposium proceedings. USDA. For. Serv. Gen. Tech. Rep. NC-1. 154 pp.nEinspahr, D. W., and M. K. Benson. 1967. Geographic variation of quaking aspen in Wisconsin and upper Michigan. Silv. Genet.16(3):106-112.nEinspahr, D. W., M. L. Harder, and M. K. Benson. 1972. Within-tree variation in fiber length of young quaking aspen. Genet. Phys. Notes. Inst. Paper Chem. Appleton, WI. No. 13.nEinspahr, D. W., J. P. Van Buijtenen, and J. R. Peckham. 1963. Natural variation and heritability in triploid aspen. Silv. Genet.12(2):51-58.nFarmer, R. E. Jr., and J. R. Wilcox. 1966. Specific gravity variation in a lower Mississippi valley cottonwood population. Tappi49(5):210-211.nIsebrands, J. 1972. The proportion of wood elements within eastern cottonwood. Wood Sci.5(2):139-146.nKennedy, R. W. 1968. Anatomy and fundamental wood properties of aspen. In J. S. Maini and J. H. Cayford, eds. Growth and utilization of poplars in Canada. Dep. For. and Rural Development. For. Branch. Dep. Publication No. 1205.nKennedy, R. W., and J. H. G. Smith. 1959. The effect of some genetic and environmental factors on wood quality in poplar. Pulp Pap. Mag. Can.59(2):37-38.nMaini, J. S., and J. H. Cayford, Editors. 1968. Growth and utilization of poplars in Canada. Can. Dep. For. Rural Dev. Publ. 1205. Ottawa. 257 pp.nPanshin, A. J., and C. DeZeeuw. 1980. Textbook of wood technology. 1. 4th ed. McGraw-Hill. New York.nPearl, I. A., and J. A. Harrocks. 1961. Studies on the chemistry of aspenwood. X. Neutral materials from the benzene extractives of Populus tremuloides. J. Org. Chem.26(5):1578-1583.nPosey, C. E., F. E. Bridgewater, and J. A. Buxton. 1969. Natural variation in specific gravity, fiber length and growth rate of eastern cottonwood in the southern great plains. Tappi52(8):1508-1511.nSteele, R. G. D., and J. H. Torrie. 1980. Principles and procedures of statistics: A biometrical approach. 2nd ed. McGraw-Hill, New York.nTaylor, F. W. 1973. Variations in the anatomical properties of South African grown Eucalyptus grandis. Appita27(3):171-178.nTaylor, F. W., and T. E. Wooten. 1973. Wood property variation of Mississippi delta hardwoods. Wood Fiber5(1):2-13.nValentine, F. A. 1962. Natural variation in Populus tremuloides in New York. 9th Northeastern For. Tree. Imp. Conf. Pp. 17-34.nVan Buijtenen, J. P., D. W. Einspahr, and P. N. Joranson. 1959. Natural variation in Populus tremuloides (Michx.). Tappi42(10):812-823.nWalters, C. S., and G. Bruckmann. 1965. Variation in specific gravity of cottonwood: as affected by tree sex and stand location. J. For.63(3):182-185.nYanchuk, A. D. 1982. Clonal variation of wood density and fibre length of trembling aspen in north-central Alberta. Unpublished M.Sc. thesis, Dep. of Forest Science, University of Alberta, Edmonton.nYanchuk, Alvin D., Bruce P. Dancik, and Michael M. Micko. 1983. Variation and heritability of wood density and fibre length of trembling aspen in Canada. Silv. Genet.32(4/5): in press.nZobel, B. 1961. The inheritance of wood properties in conifers. Silv. Genet.10(3):65-70.n
The copyright of an article published in Wood and Fiber Science is transferred to the Society of Wood Science and Technology (for U. S. Government employees: to the extent transferable), effective if and when the article is accepted for publication. This transfer grants the Society of Wood Science and Technology permission to republish all or any part of the article in any form, e.g., reprints for sale, microfiche, proceedings, etc. However, the authors reserve the following as set forth in the Copyright Law:
1. All proprietary rights other than copyright, such as patent rights.
2. The right to grant or refuse permission to third parties to republish all or part of the article or translations thereof. In the case of whole articles, such third parties must obtain Society of Wood Science and Technology written permission as well. However, the Society may grant rights with respect to Journal issues as a whole.
3. The right to use all or part of this article in future works of their own, such as lectures, press releases, reviews, text books, or reprint books.