Wood Quality Of Norway Spruce Grown In Plantations In Quebec
Keywords:
Norway spruce, Picea abies (L.) Karst, wood quality, wood density, growth rates, intraand inter-provenance variation, phenotypic selectionAbstract
This study was conducted as part of the genetic research program on Norway spruce [Picea abies (L.) Karst.] grown in Quebec. The objective of the study was to acquire more information on the wood density of Norway spruce grown in plantations and to determine whether the wood density could be used as a selection criterion for species improvement. To this end, 70-year-old commercial plantation-grown trees were first sampled in order to establish the wood density radial variation pattern and determine the existing relationships between the properties of juvenile wood and the average values of mature trees. Subsequently, the study of 22 provenances from central Europe introduced in Quebec was designed to verify the intra- and inter-provenance variation in wood density, determine the correlation between wood density and radial growth rates and origin, and estimate the impact of a selection of superior phenotypes on wood density. The study on 70-year-old trees showed that wood density declines from pith toward bark up to the eighth annual ring, stabilizes until the fourteenth annual ring, and increases thereafter. The simple regression equations calculated between the density of the juvenile wood and the mean density of the trees were significant with R2 varying from 0.53 to 0.64. The study of the wood density of the trees in the provenance trial, 19 growing seasons after planting, showed the presence of a moderate negative correlation between radial growth rate and wood density. No significant difference was found among provenances with regard to the average density values of the juvenile wood. For the radial variation pattern of wood density and annual growth ring width, comparisons made it possible to identify significant differences between southwestern and northeastern European provenances. There is still, however, strong individual variability in these traits, with 65 to 85% of total variation being due to differences between trees from the same plot. A three-stage selection procedure would make it possible to maintain wood density, to reduce by half the deviation between the internal and external juvenile wood density, and to double the volume output, depending on the selection strategy chosen. These results confirm the potential for rapid growth and the excellent wood quality of Norway spruce provenances that are well adapted to the soil and climate conditions in Quebec.References
Anonymous. 1992. Mise à l'essai des classes de bois de l'épinette de Norvège. For. Tech., Forestry Canada — Maritimes Region. 2(3): 1 and 8.nBeaulieu, J., and A. Corriveau. 1985. Variabililé de la densité du bois et de la production des provenances d'épinette blanche. 20 ans après plantation. Canadian J. Forest Res. 15:833-838.nBendtsen, B. A. 1978. Properties of wood from improved and intensively managed trees. Forest Prod. J. 28(10):61-72.nCarmirand, R., C. Camiré, and B. Bernier. 1983. Performance of Picea abies (L.) Karst. in plantation in Quebec. Canada: II. Nutrition-growth relationships. Plant Soil 73:17-26.nChang, C. I., and R. W. Kennedy. 1967. Influence of specific gravity and growth rate on dry wood production in plantation-grown white spruce. Forest. Chiron. 43(1): 165-173.nClark, A., and J. R. Saucier. 1991. Influence of planting density, intensive culture, geographic location, and species on juvenile wood formation in southern pine. Georgia Forest Research Paper, Athens, GA.nComité D'Amelioration Génétique Des Arbres Forestiers Du Québec (CAGAFQ). 1983. Amelioration génétique des essences résineuses au Québec: Recherche et developpement. A. Corriveau et G. Vallée, eds. Service canadien des forêls et ministère de l'Énergie et des Ressources du Québec, Sainte-Foy, Québec, Canada.nCorriveau, A., J. Beaulieu, and G. Daoust. 1988. Phenotypic stability and productivity of Central European Norway Spruce provenances in Quebec, Canada. IUFRO working party S 2.02-11 meeting, Tjornarp, Sweden (unpublished).nCorriveau, A., J. Beaulieu, and G. Daoust. 1991. Heritability and genetic correlations of wood characters of Upper Ottawa Valley white spruce populations grown in Quebec. Forest. Chron. 67(6):698-705.nCorriveau, A., J. Beaulieu, and F. Mothe. 1987. Wood density of natural while spruce populations in Quebec. Canadian J. Forest Res. 17:675-682.nCorriveau, A., J. Beaulieu, and F. Mothe, J. Poliquin, and J. Doucet. 1990. Densité et largeur des cernes des populations d'épinettes blanches de la région forestière des Grands Lacs et du Saint-Laurent. Canadian J. Forest Res. 20: 121-129.nDaoust, G., J. Beaulieu, and A. Corriveau. 1987. Rooting ability of cuttings from sexually mature Norway spruce. Canadian Forestry Service, Laurentian Forestry Centre, Sainte-Foy, Québec, Canada. Information Report LAU-X-75.nElliott, G. K. 1970. Wood density in conifers. Commonwealth Agricultural Bureaux, Technical Communication No. 8, Oxford, England.nJessome, A. P. 1977. Resistance et propriétes connexes des bois indigènes au Canada. Laboratoire des produits forestiers de l'Est. Rapport technique de foresterie No. 21. Ottawa, Ontario, Canada.nKeith, C. T. 1961. Characteristics of annual rings in relation to wood quality in white spruce. Forest Prod. J. 11(3):122-126.nKennedy, R. W. 1966. Intra-increment variation and heritability of specific gravity, parallel-to-grain tensile strength, stiffness, and tracheid length, in clonal Norway spruce. Tappi 49(7):292-300.nKeylwerth, R. 1954. A contribution to qualitative growth analysis. Holz Roh-Werkst. 12(3):77-83 (taken from Smith 1954).nLagercrantz, U., and N. Ryman. 1990. Genetic structure of Norway spruce (Picea abies): Concordance of morphological and allozymic variation. Evolution 44(1): 38-53.nLarson, P. R. 1969. Wood formation and the concept of wood quality. Yale University School Forest Bulletin No. 74, New Haven, CT.nMacArthur, J. D. 1964. Norway spruce plantations in Québec. Canadian Department of Forestry. Forest Research Branch. Publication No. 1059, Ottawa, Ontario, Canada.nNepveu, P., and Y. Birot. 1979. Les correlations phénotypiques juvénile-adulte pour la densité du bois et la vigueur chez l'épicéa. Annales des Sciences Forestières, 36(2):125-149.nOlesen, P. O. 1976. The interrelation between basic density and ring width of Norway spruce. Forstlige Forsogsvaesen i Danmark 34(4):339-359.nOlesen, P. O. 1977. The variation of the basic density level and tracheid width within the juvenile and mature wood of Norway spruce. Forest tree improvement, Arboretet Horsholm. Denmark, No. 12.nPanshin, A. J., and C. DeZeeuw. 1980. Textbook of wood technology, 4th ed. McGraw-Hill Book Co., New York. NY.nParadis, C. 1991. Programme de surveillance dans les plantations. Pages 14-17 in Insectes et maladies des arbres. Service de la protection contre les inseetcs et les maladies, ministère des Forêts, and Forestry Canada. Laurentian Forestry Centre, Sainte-Foy, Québec, Canada.nPolge, H. 1964. Le bois juvénile des coniferes. Revue Forestière Françhise 6:474-505.nSAS Institute Inc. 1988a. SAS user's guide: Statistics, version 6.03 edition, SAS Institute Inc., Cary, NC.nSAS Institute Inc. 1988b. SAS user's guide: Basic, version 6.03 edition. SAS Institute Inc., Cary, NC.nSell, J., and F. Kropf. 1990. Proprietes et caractéristiques des essences de bois. Lignum, Union suisse en faveur du bois. Le Mont, Suisse.nSenft, J. F., B. A. Bendtsen, and W. L. Galligan. 1985. Weak wood, fast-grown trees make problem lumber. J. Forestry 83(8):477-484.nSmith, D. M. 1954. Maximum moisture content method for determining specific gravity of small wood samples. USDA Forest Products Laboratory, Forest Science No. 2014.nWang, E. I. C., T. Mueller, and M. M. Micko. 1985. Drainage effect on growth and wood quality of some bog-grown trees in Alberta. Forest. Chron. 61(6):489-493.nWright, S. P. 1992. Adjusted P-values for simultaneous inference. Biometrics 48:1005-1013.nZobel, B. J. 1961. Inheritance of wood properties in conifers. Silvae Genetica 10(3):65-70.nZobel, B. J., R. C. Kellison, M. F. Matthias, and A. V. Hatcher. 1972. Wood density of the southern pines. Technical Bulletin No. 208. School of Forest Resources, North Carolina State University, Raleigh, NC.n
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