Influence of Stand Density on Ring Width and Wood Density at Different Sampling Heights in Black Spruce (<i>Picea Mariana</i> (Mill.) B.S.P.)

Authors

  • Jérôme Alteyrac
  • S. Y. Zhang
  • Alain Cloutier
  • Jean-Claude Ruel

Keywords:

Black spruce, stand density, sampling height, ring width, ring density, variation

Abstract

Thirty-six black spruce sample trees were collected from an 80-year-old stand to examine the influence of stand density on selected wood quality attributes and their variation with sampling height. The stand, naturally regenerated from fire in 1906, was located in Chibougamau, 400 km north of Québec. Each tree was assigned a local stand density ranging from 1390 to 3590 stems/ha, calculated from the number of neighboring trees. The trees were grouped into three stand density categories (1790, 2700, and 3400 stems/ha). Each sample tree was analyzed by X-ray densitometry, and various ring features including ring width and wood density were measured for each ring from pith to bark, at three heights (2.4, 5.1, and 7.8m) and ring area and earlywood proportion were computed. For all features studied, the variation due to sampling height was larger than that due to stand density. The longitudinal variations for ring density and earlywood density depend largely upon the wood type (juvenile wood or mature wood). A variation of ring density with sampling height in the stem from 425 to 458 kg/m3 was observed in juvenile wood, but variations with stand density in all the growth ring features studied were small. Notably, it was observed that stand density had more influence on ring width features than on ring density features.

References

Abdel-Gadir, Y. A., and R. L. Krahmer. 1993. Estimating the age of demarcation of juvenile and mature wood in Douglas-fir. Wood Fiber Sci. 25(3):242-249.nAckermann, F. 1995. The relationship between forest site and intra-ring wood density factors for Quercus robur in SW France. Annales des Sciences Forestieres 1995, 52:6, 635-652.nAlazard, P. 1994. Stand density and spacing of Pinus pinaster: Consequences for growth and tree quality. Afocel Armef No 2:129-144.nBarbour, R.J. 1987. A preliminary study of the wood properties of fast-grown black spruce, Picea mariana, from Quebec. Canadian Forest Service No. 33, Forintek Canada Corp. Ottawa, Ontario. 133 pp.nBarbour, R.J. 1991. Evaluating the influence of spacing and thinning on wood quality. Space to grow: Spacing and thinning in northern Ontario. Pages 158-162 in Proc. Symposium Sponsored by Forestry Canada, Ontario Region, and the Ontario Ministry of Natural Resources, 18-20 June 1990, Water Tower Inn, Sault Ste. Marie, Ontario. 206 pp.nBarbour, R.J. D. Sabourin, and E. Chiu. 1989. Evaluation of basic wood properties of black spruce from Quebec part II. Canadian Forest Service No. 31, Forintek Canada Corp. Ottawa, Ontario. 36 pp.nBiblis, E. J., H. F. Carino, R. Brinker, and C. W. McKee. 1995. Effect of stand density on flexural properties of lumber from two 35-year-old loblolly pine plantations. Wood Fiber Sci. 27(1):25-33.nBiblis, E. J., H. F. Carino, R. Brinker, and C. W. McKee. 1997. Flexural properties of lumber from two 40-year-old loblolly pine plantations with different stand densities. Wood Fiber Sci. 29(4): 375-380.nBues, C. T. 1985. Influence of stand density and thinning on the wood density of South African Pinus radiata. Holz Roh-Werst. 43(2): 69-73.nChui, Y. H., S. Y. Zhang, J. C. Price, and G. Chauret. 1997. Early response of balsam fir and black spruce to precommercial thinning. CTIA/IUFRO International Wood Quality Workshop, Quebec City, Canada. Timber management toward wood quality and end-product value. Chapter V pp. 15-21.nClark, A., and J. R. Saucier. 1989. Influence of initial planting density, geographic location, and species on juvenile wood formation in southern pine. Forest Prod. J. 39(7/8):42-48.nCregg, B. M., P. M. Dougherty, and T. C. Hennessey. 1988. Growth and wood quality of young loblolly pine trees in relation to stand density and climatic factors. Can. J. For. Res. 18:851-858.nDegron, R., and G. Nepveu. 1996. Predicting intra- and intertree variability of sessile oak (Quercus petraea) wood density through modelling earlywood and latewood width and density from cambial age, ring width and height within the tree. Annales des Sciences Forestieres 53 (5):1019-1030.nGartner, B. L., E. M. North, G. R. Johnson, and R. Singleton. 2002. Effects of live crown on vertical patterns of wood density and growth in Douglas-fir. Can. J. For. Res. 32:439-447.nHorvath, R. 2002. Impacts de la coupe sur la productivité des pessières noires à mousses situées dans le secteur nord-ouest de la réserve faunique Ashuapmushuan. Mémoire de maîtrise, Université Laval. 61 pp.nHuuri, O., E. Lahde, and L. Huuri. 1987. Effect of stand density on the quality and yield of young Scots pine plantations. Folia Forestalia, Institutum Forestale Fenniae No 685. 48 pp.nKellison, R. C., R. Lea, and D. J. Frederick. 1983. Effect of silvicultural practices on wood quality of southern hardwoods. Tappi J. 66(1):67-69nKoga, S., and S. Y. Zhang. 2002. Relationships between wood density and annual growth rate components in Balsam fir (Abies balsamea). Wood Fiber Sci. 34(1): 146-157.nKoga, S., J. Tsutsumi, and K. Oda. 1995. Juvenile wood percentages of karamatsu (Larix leptolepis). Bulletin of the Kyushu University Forests 72:217-227.nKoubaa, A., S. Y. Zhang, N. Isabel, J. Beaulieu, and J. Bousquet. 2000. Phenotypic correlations between juvenile-mature wood density and growth in black spruce. Wood Fiber Sci. 32(1):61-71.nKoubaa, A., S. Y. Zhang, S. Makni, C. Plomion, and C. Cahalan. 2002. Defining the transition from earlywood to latewood in black spruce based on intra-ring wood density profiles from X-ray densitometry. Annals of Forest Science 59: 5/6, 511-518.nKromhout, C. P. 1968. The influence of stand density on certain anatomical wood properties of Pinus pinaster in two C. C. T. stands. Bosb. Suid-Afr. 9:45-66.nLaw, K. N., and J. L. Valade. 1997. Effect of wood quality on thermomechanical pulping: A case study on black spruce and jack pine. CTIA/IUFRO International Wood Quality Workshop, Quebec City, Canada. Timber management toward wood quality and end-product value. Chapter VII pp. 3-7.nLei, H., B. L. Gartner, and M. R. Milota. 1997. Effect of growth rate on the anatomy, specific gravity, and bending properties of wood from 7-year-old red alder (Alnus rubra). Can. J. For. Res. 27:80-85.nMiddleton, G. R., L. A. Jozsa, L. C. Paka, B. D. Munro, and P. Sen. 1995. Lodgepole pine product yield related to differences in stand density. Forintek, SP 35. 66 pp.nPanshin, A. J., and C. de Zeeuw. 1980. Textbook of wood technology. McGraw-Hill Book Co., New York, NY. 772 pp.nRaulier, F., C.-H. Ung, and D. Ouellet. 1996. Influence of social status on crown geometry and volume increment in regular and irregular black spruce stands. Can. J. For. Res. 26:1742-1753.nSauter, U. H., R. Mutz, and B. D. Munro. 1999. Determining juvenile-mature wood transition in Scots pine using latewood density. Wood Fiber Sci. 31(4):416-425.nUng, C.-H., F. Raulier, D. Ouellet, and J.-F. Dhote. 1997. L'indice de compétition interindividuelle de Schütz. Can. J. For. Res. 27:521-526.nWent, F. W. 1942. Some physiological factors in the aging of a tree. Pages 330-334 in Proc. 9th West. Shade Tree Conf. California Institute of Technology, Pasadena, Calif.nYang, K. C., C. A. Benson, and J. K. Wong. 1986. Distribution of juvenile wood in two stems of Larix laricina. Can. J. Forest Res. 16:1041-1049.nZhang, S. Y. 1998. Effect of age on the variation, correlations and inheritance of selected wood characteristics in black spruce (Picea mariana). Wood Sci. Technol. 32:197-204.nZhang, S. Y. and E. K. Morgenstern. 1995. Genetic variation and inheritance of wood density in black spruce (Picea mariana) and its relationship with growth: implication for tree breeding. Wood Sci. Technol. 30(1):63-75.nZhang, S. Y. and G. Chauret. 2001. Impact of initial spacing on tree and wood characteristics, product quality and value recovery in black spruce (Picea mariana). Canadian Forest Service Report. No. 35, Forintek Canada Corp., Sainte-Foy, Quebec. 47 pp.nZhang, S. Y. R. Gosselin, and G. Chauret. 1997. Wood quality: Its definition, impact, and implications for value-added timber management and end uses. Proc. of the CTIA-IUFRO International Wood Quality Workshop, Quebec City, Canada, August 18-22. Pp. 1.17-1.39n

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Published

2007-06-05

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Research Contributions