Pits as Natural Irregularities in Softwood Fibers
Keywords:
Pit size, pit shape, pit orientation, pit density, fiber strength, <i>Pinus silvestris</i>, <i>Picea abies</i>Abstract
Parameters for pit size, shape, orientation, and density, and their correlations with fiber dimensions in Scots pine and Norway spruce fibers were determined. Bordered pits were smaller with greater cross-sectional fiber area, and more circular the wider the fibers. Angular pit orientation became more transverse in pine fibers, and pit density increased in both species with fiber width. Normalized pit size and pit density increased towards fiber tips, especially in thin-walled pine fibers. These observations can be explained by the developing process and the functional role of tracheids in wood. The appearance of pits as stress-enhancing irregularities in fiber structure should be considered in fiber-network theories predicting paper properties, as well as in the measurement of the mechanical properties of fibers.References
Barnett, J. R. 1981. Secondary xylem cell development. Pages 47-95 in J. R. Barnett, ed. Xylem cell development. Castle House Publications Ltd., Tunbridge Wells, Kent, UK.nBarnett, J. R., and J. M. Harris. 1975. Early stages of bordered pit formation in radiata pine. Wood Sci. Technol. 9(3): 233-241.nBernhart, A. 1964. Über die Rohdichte von Fichtenholz. On the specific gravity of the wood of Norway spruce (Picea abies (L.) Karst.). Holz Roh- Werkst. 22(6):215-228.nBisset, I. J. W., and H. E. Dadswell. 1950. The variation in cell length within one growth ring of certain angio-sperms and gymnosperms. Austr. For. 14(1): 17-29.nBrown, C. L. 1970. Physiology of wood formation of conifers. Wood Sci. 3(1):8-22.nCox, H. L. 1952. The elasticity and strength of paper and other fibrous materials. Br. J. Appl. Phys. 3:72-79.nEl-Hosseiny, F., and D. H. Page. 1975. The mechanical properties of single wood-pulp fibers: theories of strength. Fiber Sci. Technol. 8:21-30.nEvans, R. 1994. Rapid measurement of the transverse dimensions of tracheids in radial wood sections from Pinus radiata. Holzforschung 48(2): 168-172.nFengel, D., and M. Stoll. 1973. Über die Veränderungen des Zellquerschnitts, der Dicke der Zellwand und der Wandschichten von Fichtenholz-Tracheiden innerhalb eines Jahrringes. On the variation of the cell cross area, the thickness of the cell wall and of the wall layers of sprucewood tracheids within an annual ring. Holzforschung 27(1): 1-7.nGroom, L. H., S. M. Shaler, and L. Mott. 1995. Characterizing micro- and macromechanical properties of single wood fibers. Pages 13-22 in Sept. 11-14, 1995 Paper Physics Conference, Niagara-on-the-lake, Ontario, Canada.nHelle, T. 1963. Some aspects on fibre strength and fibre bondings in sulphate and sulphite paper. Sven. Papperstidn. 66(24):1015-1030.nJohansson, D. 1939. Något om vår- och höstved hos tall och gran och dess inverkan på sulfit- och sulfatmassans egenskaper. Summary: On the spring and summer wood of pine and spruce and their influence on the properties of sulphite and sulphate pulp. Finn. Pap. Timber J, 7a (special number): 54-72.nKoran, Z. 1974. Intertracheid pitting in radial walls of black spruce tracheids. Wood Sci. 7(2):111-115.nKoran, Z. 1977. Tangential pitting in black spruce tracheid. Wood Sci. Technol. 11(2):115-123.nKozlowski, T. T. 1971. Growth and development of trees, vol. II. Cambial growth, root growth and reproductive growth. Academic Press, New York, NY. 514 pp.nLiese, W. 1963. The fine structure of bordered pits in softwoods. Pages 271-290 in W. A. Côté Jr., ed. Cellular ultrastructure of woody plants. Syracuse, NY.nLin, J. 1989. Distribution, size and effective aperture area of the inter-tracheid pits in the radial wall of Pinus radiata tracheids. IAWA Bulletin n.s. 10(1):53-58.nOkamura, S., H. Saiki, and H. Harada. 1973. Polarizing microscope study on the concentric orientation of nγ in the pit border region of softwood tracheids. Holzforschung 27(1):12-16.nOkamura, S., H. Harada, and H. Saiki. 1974. The variation in the cell wall thickness along length of a conifer tracheid. Bull. Kyoto Univ. For. 46:162-169.nPage, D. H., and E El-Hosseiny. 1983. The mechanical properties of single wood pulp fibers. Part VI. Fibril angle and the shape of the stress-strain curve. J. Pulp Paper Sci. 84(9):TR99-100.nPage, D. H., E. El-Hosseiny., K. Winkler, and A. P. S. Lancaster. 1977. Elastic modulus of single wood pulp fibers. Tappi 60(4):114-117.nPage, D. H., R. S. Seth, and E El-Hosseiny. 1985. Strength and chemical composition of wood pulp fibers. Pages 77-91 in Eighth Fundamental Research Symposium, 1985. Oxford UK.nTakizawa, T. 1974. The distribution of pits on one tracheid of Karamatsu (Larix leptolepis Gord.). Res. Bull. Coll. Exp. For., Hokkaido Univ. 31:481-500.nTakizawa, T., and S. Ishida. 1972. The number and distribution of pits on one tracheid, and their variation in a trunk of Todomatsu (Abies sachalinensis Fr. Schun.). Res. Bull. Coll. Exp. For., Hokkaido Univ. 29:189-205.nThomas, R. J., and J. L. Scheld. 1967. The distribution and size of the inter-tracheid pits in an eastern hemlock. Forest Sci. 13:85-89.nWardrop, A. B., and H. E. Dadswell. 1957. Variations in the cell wall organization of tracheids and fibres. Holzforschung 11(2):33-41.n
Downloads
Published
Issue
Section
License
The copyright of an article published in Wood and Fiber Science is transferred to the Society of Wood Science and Technology (for U. S. Government employees: to the extent transferable), effective if and when the article is accepted for publication. This transfer grants the Society of Wood Science and Technology permission to republish all or any part of the article in any form, e.g., reprints for sale, microfiche, proceedings, etc. However, the authors reserve the following as set forth in the Copyright Law:
1. All proprietary rights other than copyright, such as patent rights.
2. The right to grant or refuse permission to third parties to republish all or part of the article or translations thereof. In the case of whole articles, such third parties must obtain Society of Wood Science and Technology written permission as well. However, the Society may grant rights with respect to Journal issues as a whole.
3. The right to use all or part of this article in future works of their own, such as lectures, press releases, reviews, text books, or reprint books.