Variation in the Relationship of Wood Density With Growth In 40 Black Spruce (<i>Picea Mariana</i>) Families Grown in New Brunswick
Keywords:
Black spruce, wood density, growth traits, dry mass weight, variation, correlation, family, environmentAbstract
This study examined variation in the relationship of wood density with growth traits (DBH, tree height, and bole volume) in 15-year-old half-sib families of black spruce (Picea mariana) grown on two different sites located in northern New Brunswick. Effect of genotype and environment on wood density was also briefly discussed. Although a moderately negative relationship between wood density and growth generally exists, the relationship, to some extent, varies with genotype and environment. A nonsignificant or even a weak positive relationship between wood density and growth can be found in some families. Moreover, the negative relationship between wood density and growth appears to be weaker in families growing in a more favorable environment. This study indicates that it is possible to select some families that not only grow fast, but also maintain high wood density.References
Abdel-Gadir, A. Y., R. L. Krahmer, and M. D. Mckimmy. 1993. Intra-ring variation in mature Douglas-fir trees from provenance plantations. Wood Fiber Sci. 25(2): 170-181.nAllen, P. J. 1977. Genotypic and phenotypic correlations of wood and tree characteristics in: Variation of growth, stem, quality, and wood properties in relation to genetic improvement of tropical forest trees. IUFRO Workshop, Brisbane, Australia.nAnonymous. 1978. Black spruce seedling seed orchard and progeny tests. New Brunswick Tree Improv. Council Estab. Rep. No. 3.nBendtsen, B. A. 1978. Properties of wood from improved and intensively managed trees. Forest Prod. J. 28:61-72.nCanada Department of Forestry. 1963. Native trees of Canada, 6th ed. Bulletin 61, Roger Duhamel, Ottawa, Canada.nDorn, D. 1969. Relationship of specific gravity and tracheid length to growth rate and provenance in Scotch pine. 16th Northeastern For. Tree Improv. Conf., Québec, Canada.nFalconer, D. S. 1981. Introduction to quantitative genetics. Ronald Press, New York, NY.nHall, J. P. 1984. The relationship between wood density and growth rate and the implications for the selection of black spruce plus trees. Inf. Rep. N-X-224, Newfoundland For. Res. Cent., Can. For. Serv., Newfoundland, Canada.nHoner, T. G., M. F. Ker, and I. S. Alemdag. 1983. Metric timber tables for the commercial tree species of Central and Eastern Canada. Inf. Rep. M-X-140, Marit. For. Res. Cent., Fredericton, New Brunswick, Canada.nJourdain, C. J., and J. R. Olson. 1984. Wood property variation among forty-eight families of American sycamore. Wood Fiber Sci. 16(4):498-507.nKeith, C. T., and R. M. Kellogg. 1986. Wood quality considerations in tree improvement programs. Proc. Workshop held in Quebec City, August 19, 1995. Special publication SP-513E, Forintek Canada Corp., Ottawa, Canada.nKellogg, R. M. 1989. Second growth Douglas-fir: Its management and conversion for value. Special Publication No. SP-32, Forintek Canada Corp., Vancouver, B.C., Canada.nMagnussen, S., and C. T. Keith. 1990. Genetic improvement of volume and wood properties of jack pine: Selection strategies. For. Chron. 66:281-286.nMckimmy, M. D. 1966. A variation and heritability study of wood specific gravity in 46-year-old Douglas-fir from known seed sources. Tappi 49(2):542-549.nMckimmy, M. D., and D. D. Nicholas. 1971. Genetic differences in wood traits among half-century-old families of Douglas-fir. Wood Fiber 2(4):347-355.nMorgenstern, K. E., and M. Villeneuve. 1986. The role of wood quality in New Brunswick's genetic improvement program. XVII IUFRO World Congress, Ljubljana, Yugoslavia.nMullins, E. J., and T. S. Mcknight. 1981. Canadian woods: Their properties and uses, 3rd ed. Univ. Toronto Press, Toronto, Canada.nPanshin, A. J., and C. de Zeeuw. 1980. Textbook of wood technology, 4th ed. McGraw-Hill Book Company, New York, NY.nPark, Y. S., J. D. Simpson, D. P. Fowler, and E. K. Morgenstern. 1989. Selection index with desired gains to rogue jack pine seedling seed orchard. Inf. Rep. M-X-176, For. Can. Marit. Reg., Fredericton, New Brunswick.nRowe, J. S. 1972. Forest regions of Canada. Can. For. Serv. Publ. 1300. Ottawa, Canada.nShelbourne, C. J., I.J. Thulin, and R. M. Scott. 1972. Variation, inheritance, and correlation amongst growth, morphological, and wood characters in radiata pine. New Zealand For. Serv., Prod. For. Div. Gen. and Tree Improv. Rep. 61.nSmyth, J. H., and A. J. Brownright. 1986. Forest tree production centres in Canada—1984. Inf. Rep. O-X-378, Can. For. Serv. Great Lakes For. Res. Cent. Sault Ste. Marie, Ontario, Canada.nSohn, S. I., and R. E. Goddard. 1974. Genetic study of wood specific gravity in slash pine. 2nd Northeastern For. Tree Improv. Conf., Syracuse, NY.nSpurr, S. H., and W. Hsuing. 1954. Growth rate and specific gravity in conifers. J. Forestry 52:192-200.nSquillace, A. E., R. M. Echols, and K. M. Dorman. 1962. Heritabilities of specific gravity and summer-wood percent and relation to other factors in slash pine. Tappi 45:599-601.nVargas-Hernandez, J., and W. T. Adams. 1991. Genetic variation of wood density components in young coastal Douglas-fir: Implications for tree breeding. Can. J. For. Res. 21:1801-1807.nWeiner, J., and L. Roth. 1966. The influence of environment and genetics on pulpwood quality. Bibliographic Series No. 224, Inst. Paper Chem., Appleton, WI.nYanchuk, A. D., and G. K. Kiss. 1993. Genetic variation in growth and wood specific gravity and its utility in the improvement of interior spruce in British Columbia. Silvae Genet. 42(2-3): 141-147.nYanchuk, A. D., B. P. Dancik, and M. M. Micko. 1984. Variation and heritabilities of wood density and fibre length of trembling aspen in Alberta, Canada. Silvae Genet. 33:11-16.nZhang, S. Y. 1992. Structure-property relationship of wood in East-Liaoning oak. Wood Sci. Technol. 26:139-149.nZhang, S. Y. 1994. Mechanical properties in relation to specific gravity in 342 Chinese woods. Wood Fiber Sci. 26(4):512-526.nZhang, S. Y. 1995a. Effect of growth rate on wood specific gravity and selected mechanical properties in individual species from distinct wood categories. Wood Sci. Technol. 29(6):451-465.nZhang, S. Y. 1995b. The variation, correlations and inheritance of intraring wood density and width in black spruce (Picea mariana) trees from 15-year-old families. (in preparation).nZhang, S. Y., and E. K. Morgenstern. 1996. Genetic variation and inheritance of wood density in black spruce (Picea mariana) and its relationships with growth: Implication for breeding. Wood Sci. Technol. (in press).nZhang, S. Y., and Y. Zhong. 1991. Effect of growth rate on specific gravity of East-Liaoning oak (Quercus liaotungensis) wood. Can. J. For. Res. 21:255-260.nZhang, S. Y., S. Fujiwara, and P. Baas. 1992. Hardwood density in relation to fibre morphology and tissue proportions. Proc. IUFRO All-Division 5 Conference, 23-28 August 1992, Nancy, France.nZhang, S. Y., G. Nepveu, and R. Eyono Owoundi. 1994. Intratree and intertree variation in selected wood quality characteristics of European oak (Quercus petraea and Quercus robur). Can. J. For. Res. 24:1818-1823.nZobel, B. J., and J. P. Van Buijtenen. 1989. Wood variation: Its causes and control. Springer-Verlag, Berlin, Germany.nZobel, B. J., E. Thorbjornsen, and E. Henson. 1960. Geographic, site and individual tree variation in the wood properties of loblolly pine. Silvae Genet. 9:149-158.n
Downloads
Published
Issue
Section
License
The copyright of an article published in Wood and Fiber Science is transferred to the Society of Wood Science and Technology (for U. S. Government employees: to the extent transferable), effective if and when the article is accepted for publication. This transfer grants the Society of Wood Science and Technology permission to republish all or any part of the article in any form, e.g., reprints for sale, microfiche, proceedings, etc. However, the authors reserve the following as set forth in the Copyright Law:
1. All proprietary rights other than copyright, such as patent rights.
2. The right to grant or refuse permission to third parties to republish all or part of the article or translations thereof. In the case of whole articles, such third parties must obtain Society of Wood Science and Technology written permission as well. However, the Society may grant rights with respect to Journal issues as a whole.
3. The right to use all or part of this article in future works of their own, such as lectures, press releases, reviews, text books, or reprint books.