Effects of Plantation Density on Wood Density and Anatomical Properties of Red Pine (<i>Pinus Resinosa</i> AIT.)

Authors

  • J. Y. Zhu
  • C. Tim Scott
  • Karen L. Scallon
  • Gary C. Myers

Keywords:

Red pine, small-diameter trees, forest thinning, plantation density, pulp yield, tracheid (cell) geometry, SilviScan, maceration, radial growth, latewood content

Abstract

This study demonstrated that average ring width (or average annual radial growth rate) is a reliable parameter to quantify the effects of tree plantation density (growth suppression) on wood density and tracheid anatomical properties. The average ring width successfully correlated wood density and tracheid anatomical properties of red pines (Pinus resinosa Ait.) from a never-thinned experimental plantation forest with five different initial plantation densities. The results indicate that plantation density has a pronounced effect on earlywood growth, which resulted in increased latewood volumetric fraction and uniform tracheid radial and wall thickness distribution. A similar approach of average ring width correlations may be applied to study wood and tracheid properties of trees growing in the densely populated natural forests.

References

Ballard L. A., and J. N. Long. 1988. Influence of stand density on log quality of lodgepole pine. Can. J. For. Res.18:911-916.nDinwoodie J. M. 1965. The relationship between fiber morphology and paper properties: A review of literature. TAPPI J.48(8):440-447.nDutilleul P., M. Herman, and T. Avella-Shaw. 1998. Growth rate effects on correlations among ring width, wood density, and mean tracheid length in Norway spruce (Picea abies). Can. J. For. Res.28:56-68.nEvans, R. 1994. Rapid measurement of the transverse dimensions of tracheids in radial wood sections from Pinus radiata, Holzforschung48(2):168-172.nEvans, R., G. Downes, D. Menz, and S. Stringer. 1995. Rapid measurement of variation in tracheid transverse dimensions in a radiata pine tree. Appita48(2):134-138.nHatton, J. V., K. Hunt, and W. Y. Gee. 1996. Effects of spacing on the basic relative density, wood and fiber properties of plantation grown western cedar. Pulp and Paper Research Institute of Canada, Vancouver, B.C. Pulp and Paper Rep. 1239.nJanas, P. S., and D. G. Brand. 1988. Comparative growth and development of planted and natural stands of jack pine. Forest Chronicles64(4):320-328.nKang K-Y, S. Y. Zhang, and S. D. Mansfield. 2004. The effects of initial spacing on wood density, fibre and pulp properties in jack pine (Pinus Banksiana Lamb.). Holzforschung58:455-463.nKlungness, J. H., R. Gleisner, D. Mann, K.L. Scallon, J. Y. Zhu, E.G. Horn, and L. L. Edwards. 2006. Evaluation of forest thinning materials for TMP production. TAPPI J.5(4):17-22.nKoch, P. 1972. Utilization of the Southern Pines, Vol. I The Raw Material. U.S. Dept. of Agriculture Forest Service, US Government Printing Office, Stock No: 0100-1483, Washington, DC 20402, Library of Congress Card No: 79-188-856. Pp 68-75.nLarocque, G. R. 2002. Examining different concepts for the development of a distance-dependent competition model for red pine diameter growth using long-term stand data differing in initial stand density. Forest Sci.48:24-34.nLarson, P. R. 1963. The indirect effect of drought on tracheid diameter in red pine. Forest Sci.9(1):52-62.nLindstrom, H. 1996. Basic density in Norway spruce. Part III. Development from pith outwards. Wood Fiber Sci.28(4):391-405.nMaeglin, R. R. 1967. Effect of tree spacing on weight yields for red pine and jack pine. J. Forestry65:647-650.nMakinen, H., P. Saranpaa, and S. Linder. 2002. Wood-density variation of Norway spruce in relation to nutrient optimization and fiber dimensions. Can. J. For. Res.32: 185-194nMyers, G. C. 2004. Optimizing lodgepole pine submerchantable log thermomechanical pulp. Wood Fiber Sci.36(4):476-482.nMyers, G. C., R. J. Barbour, and S. Abubakr. 1999. Small-diameter trees used for thermomechanical pulps. TAPPI J.82(10):105-110.nMyers, G. C., R. J. Barbour, and S. Abubakr. 2003. Small-diameter trees used for chemithermo-mechanical pulps. FPL-GTR-141. USDA Forest Service, Forest Products Laboratory, Madison, WI. 12 pp.nSeth, R. S. 1990. Fibre quality factors in papermaking-II. The importance of fibre coarseness, in MRS Symposium Proceedings, Materials Research Society, Pittsburgh, PA. Vol. 197, Pp 143-161.nTurnblom, E. C., and T. E. Burk. 2000. Modeling self-thinning of unthinned Lake States red pine stands using nonlinear simultaneous differential equations. Can. J. For. Res.30:1410-1418.nVahey, D. W., J. Y. Zhu, and C. T. Scott. 2007. Wood density and anatomical properties in suppressed-growth trees: Comparison of two methods. Wood Fiber Sci.39(3):462-471.nWatson P., C. Garner, R. Robertson, S. Reath, W. Gee, and K. Hunt. 2003. The effects of initial tree spacing on the fibre properties of plantation-grown coastal western hemlock. Can. J. For. Res.33:2460-2468.nYang, K. C., and G. Hazenberg. 1994. Impact of spacing on tracheid length, relative density, and growth rate of juvenile wood and mature wood in Picea mariana.Can. J. For. Res.24:996-1007.nZahner, R., and W. W. Oliver. 1962. The influence of thinning and pruning on the date of summer wood initiation in red and jack pines. Forest Sci.8(1):51-63.nZahner, R., J.E. Lotan, and W. D. Baughman. 1964. Early-wood-latewood features of red pine grown under stimulated drought and irrigation. Forest Sci.10(3):361-370.nZhu, J. Y., and G. C. Myers. 2006. Effect of plantation density on kraft pulp production from red pine (Pinus resinosa Ait.), J. Pulp Pap. Sci.32(3):187-193.nYang, K. C., C.A. Benson, and J. K. Wong. 1986. Distribution of juvenile wood in two stems of Larix laricina.Can. J. For. Res.16:1041-1049.nYeung, E. C. (1998) A beginner's guide to the study of plant structure, in Tested Studies for Laboratory Teaching, Volume 19. Karcher, S. J., Ed. Proceedings of the 19th Workshop/Conference of the Association for Biology Laboratory Education (ABLE). Pp. 125-141; also http://www.zoo.utoronto.ca/able/volumes/vol-19/09-yeung/09-yeung.htm#Maceration%20procedure.n

Published

2007-09-27

Issue

Section

Research Contributions