Intra-Annual Cellular Characteristics and their Implications for Modeling Softwood Density
Keywords:Wood density, latewood percentage, anatomy, model, Pinus sylvestris, earlywood, variability, wood quality
AbstractA numeric wood density model is presented to investigate the potential and actual influence of intra-annual cell characteristics on wood density. The model was designed for conifers with a distinctive early-latewood transition, and its performance was tested with data from Scots pine trees. The model is a helpful tool to understand how and to what extent cell characteristics influence wood density. Latewood percentage proved to be an excellent predictor for wood density, and the interaction of density with cell characteristics is shown.
Abdel-Gadir, A. Y., R. L. Krahmer, and M. D. Mc-Kimmy. 1993. Relationships between intra-ring variables in mature Douglas-fir trees from provenance plantations. Wood Fiber Sci. 25(2):182-191.nDe Kort, I. 1993a. Wood production and latewood percentage of Douglas-fir from different stands and vitality classes. Can. J. For. Res. 23:1480-1486.nDe Kort, I. 1993b. Relationships between sapwood amount, latewood percentage, moisture content, and crown vitality of Douglas fir, Pseudotsuga menziesii. IAWA J. 14(4): 413-427.nDenne, M. P. 1989. Definition of latewood according to Mork (1928). IAWA Bull. n.s. 10(1):59-62.nGilmore, A. R., G. E. Metcalf, and W. R. Boggess 1959. Specific gravity of shortleaf pine and loblolly pine in southern Illinois. J. Forestry 59:894-896.nGoggans, J. F. 1964. Correlation and heritability of certain wood properties in loblolly pine (Pinus taeda L.). Tappi 47(6):318-322.nJackson, L. W., and R. E. Strickland. 1962. Geographic variation in tracheid length and wood density of loblolly pine. Georgia Res. Counc. 8:1-4.nKellogg, R. M., and F. F. Wangaard. 1969. Variation in the cell-wall density of wood. Wood Fiber 1(3): 180-204.nKollmann, F. F. P., and W. A. Côtê, Jr. 1968. Principles of wood science and technology. Vol. 1. Solid wood. Springer-Verlag, New York, NY.nLadell, J. T. 1959. A new method of measuring tracheid length. Forestry 32:124-125.nLarson, P. R. 1973. The physiological basis for wood specific gravity in conifers. IUFRO Div 5 Meeting, Brisbane, Australia 2:672-680.nLeclerq, A. 1980. Relationships between beechwood anatomy and its physico-mechanical properties. IAWA Bull.n.s. 1(1-2):65-71.nMatziris, D. I., and B. J. Zobel. 1973. Inheritance and correlation of juvenile characteristics in loblolly pine (P. taeda). Silvae Genet. 22:104-106.nPanshin, A. J., and C. De Zeeuw. 1980. Textbook of wood technology. 4th ed. McGraw-Hill Book Company, New York, NY.nQuirk, T. 1984. Shrinkage and related properties of Douglas-fir cell walls. Wood Fiber Sci. 16(1):115-133.nReck, S. 1963. Die Variation und Kovariation einiger anatomischer Holzmerkmale und ihre Beziehung zur Raumdichte bei Fichtenholz. Ph.D. thesis, University of Hamburg, Germany.nRoelofsen, P. A. 1959. The plant cell wall. Gebrueder Bortraeger, Berlin, Germany.nSaka, S., and R. J. Thomas. 1982. A study of lignification on loblolly pine tracheids by the SEM-EDXA technique. Wood Sci. Technol. 16:1-18.nSell, J. 1989. Eigenschaften und Kenngroessen von Holzarten. Bauverlag, Zurich, Switzerland.nSmith, D. M. 1965. Rapid measurement of tracheid cross-sectional dimensions of conifers. Forest Prod. J. 15(8): 325-334.nTrendelenburg, R., and H. Mayer-Wegelin. 1955. Das Holz als Rohstoff. 2th ed. Carl Hauser Verlag, Muenchen, Germany.nTsoumis, G. 1964. Microscopic measurement of the amount of cell wall substance in wood and its relationship to specific gravity. Tappi 47(11):675-677.nVan Buijtenen, J. P. 1964. Anatomical features influencing wood specific gravity of slash pines and the implications for the development of a high-quality pulpwood. Tappi 47:401-404.nWilson, B. F., and R. A. Howard. 1968. A computer model for cambial activity. Forest Sci. 14(1):77-90.nWimmer, R. 1991. Beziehungen zwischen Jahrringpar-ametern und Rohdichte bei Kiefernholz. Holzforsch. Holzverwert. 43(4):79-82.nWodzicki, T. J. 1971. Mechanism of xylem differentiation in Pinus silvestris L. J. Exp. Bot. 22(72):670-687.nYao, J. 1970. Influence of growth rate on specific gravity and other selected properties on loblolly pine. Wood Sci. Technol. 4:163-175.nZhang, S. Y., and Y. Zhong. 1992. Structure-property relationship of wood in East-Liaoning oak. Wood Sci. Technol. 26:139-149.nZhang, S. Y., R. Eyeno, G. Nepveu, F. Mothe, and J.-F. Dhote. 1993. Modelling wood density in European oak (Quercus petraea and Quercus robur) and simulating the silvicultural influence. Can. J. For. Res. 23:2587-2593.nZobel, B. J., and J. P. Van Buijtenen. 1989. Wood variation-Its causes and control. Springer Series in Wood Science, Springer-Verlag, New York, NY.n
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