Relationships Between Wood Density and Annual Growth Rate Components in Balsam Fir (<i>Abies Balsamea</i>)
Keywords:
Balsam fir, wood density, ring width, earlywood, latewood, juvenile wood, mature wood, variation, correlationAbstract
This study examined relationships of wood density components with annual growth rate components (or annual ring width components) in juvenile wood and mature wood of balsam fir [Abies balsamea (L.) Mill.]. The relationships were studied at two different levels: 1) inter-tree level (between trees), and 2) intra-tree level (within a tree). In addition, juvenile-mature wood correlations for these characteristics were investigated. Wood density and annual ring width components of individual growth rings were measured by X-ray densitometry. Based on tree averages (at the inter-tree level), wood density is significantly correlated with its components (earlywood density, latewood density) and latewood percentage in both juvenile wood and mature wood; and earlywood density and latewood percentage are the two most important parameters in determining the overall wood density of the tree. Wood density, however, is not significantly correlated with annual growth rate (ring width) in either juvenile wood or mature wood, although a weakly negative correlation tends to strengthen in mature wood. This suggests that the relationship between wood density and annual growth rate in this species may vary with cambial age. Intra-ring wood density variation (IDV) shows a positive correlation with wood density traits, latewood width, and latewood percentage in both juvenile wood and mature wood, whereas a weakly negative correlation of IDV with ring width and earlywood width exists in balsam fir. Latewood traits are the most important parameters in determining the intra-ring wood density uniformity. At the intra-tree level (based on ring averages within a tree), relationships between wood density components and ring width components are similar to those found between the trees, although some relationships, to some extent, vary with tree. For each wood density trait, the juvenile-mature wood correlation is significant but moderate. For this species, earlywood density in juvenile wood seems to be the best parameter for predicting mature wood density.References
Abdel-Gadir, A. Y., R. L. Krahmer, and M. D. McKimmy. 1993. Relationships between intra-ring variables in mature Douglas-fir trees from provenance plantation. Wood Fiber Sci.25(23): 182-191.nBarbour, R. J., R. E. Bailey, and J. A. Cook. 1992. Evaluation of relative density, diameter growth, and stem form in a red spruce (Picea rubes) stand 15 years after precommercial thinning. Can. J. For. Res.22:229-238.nBlouin, D., J. Beaulieu, G. Daoust, and J. Poliquin. 1994. Wood quality of Norway spruce grown in plantations in Quebec. Wood Fiber Sci.26(1):61-71.nCorriveau, A., J. Beaulieu, and F. Mothe. 1987. Wood density of natural white spruce populations in Quebec. Can. J. For. Res.17(7):675-682.nDorn, D. 1969. Relationship of specific gravity and tra-cheid length to growth rate and provenance in Scots pine. 16th Northeastern For. Tree Improv. Conf., Quebec City, Quebec.nHall, J. P. 1984. The relationship between wood density and growth rate and the implications for the selection of black spruce. Newfoundland For. Res. Cent. Inf. Rep. N-X-224, Can. For. Serv., St-John's, Newfoundland. 22 pp.nJozsa, L. A., J. E. Richard, and S. G. Jonson. 1987. Calibration of Forintek's direct reading X-ray densitometer. Forintek Canada Corp., CFS Report No. 36a, Vancouver, BC.nJessome, A. P. 2000. Strength and related properties of woods grown in Canada. Forinteck Canada Corp., Publication SP-514E, Sainte-Foy, Quebec. 37 pp.nJourdain, C. J., and J. R. Olson. 1984. Wood property variation among forty-eight families of American sycamore. Wood Fiber Sci.16(4):498-507.nKeith, C. T. 1961. Characteristics of annual rings in relation to wood quality for white spruce. Forest Prod. J.11(3): 122-126.nKeller, R., and M. Thoby. 1977. Liaisons entre l'état juvénile et l'état adulte pour quelqes caractères technologiques et auxométriques chez le Douglas (Pseudot-suga menziesii). Ann. Sci. For.34(3): 175-203.nKennedy. R. W. 1995. Coniferous wood quality in the future: concerns and strategies. Wood Sci. Technol.29: 321-338.nKoga. S., K. Oda, J. Tsutsumi, and T. Fujimoto. 1996. Effect of thinning on basic density and tracheid length of karamatsu (Larix leptolepis). Mokuzai Gakkaishi42(6):605-611.nKoga. S., K. Oda, J. Tsutsumi, and T. Fujimoto. 1997. Variation in basic wood properties of sugi (Cryptomeria japonica) Culti-vars. Pages VIII-11-16 in S. Y. Zhang, R. Grosselin, and G. Chauret, eds. Timber management toward wood quality and end-product value, Proc. CTIA/IUFRO International Wood Quality Workshop, Quebec City, Quebec, 550 pp.nKoga. S., S. Y., Zhang, and J. Begén. 2001. Effects of precommercial thinning on annual growth and wood density in balsam (Abies balsamea). Wood Fiber Sci. (submitted)nKoubaa, A., S. Y. Zhang, N. Isabel, J. Beaulieu, and J. Bousquet. 2000. Phenotypic correlations between juvenile-mature wood density and growth in black spruce. Wood Fiber Sci.32(1):61-71.nMagnussen, S., and C. T. Keith. 1990. Genetic improvement of volume and wood properties of jack pine. Selection strategies. For. Chron.66:281-286.nMullins, E. J., and T. S. McKnight. 1981. Canadian woods: Their properties and uses. 3rd ed., University of Toronto, Toronto, Ontario. 389 pp.nNepveu, G., and Y. Birot. 1979. Les corrélations phénotypiques juvénile-adulte pour la densité du bois et de la vigueur chez l'épicea. Ann. Sci. For.36(2):125-149.nPark, Y. S. J., D. Simpson, D. P. Fowler and E. K. Morgenstern. 1989. Selection index with desired gains to rogue jack pine seedling seed orchard. Inf. Rep. M-X-176, For. Can. Marit. Reg., Fredericton, New Brunswick.nPanshin, A. J., and C. De Zeeuw. 1980. Textbook of wood technology. 4th ed. McGraw-Hill Book Company, New York, NY.nSAS Institute Inc. 1988. SAS/STAT user's guide. Release 6.03 ed. SAS Institute Inc., Cary, NC.nSchneider, R. 2001. Effet d'une éclaircie précommercial 28 ans aprés traitement sur la productivité d'une sapinière en Gaspésie. Mémoire de maítrise, Faculté de foresterie et de géomatique. Université Laval, Québec. 31 pp.nVargas-Hernandez, J., and W. T. Adams. 1991. Genetic variation of wood density components in young coastal Douglas-fir: Implication for tree breeding. Can. J. For. Res.21:1801-1807.nZarnovican, R., and C. Laberge. 1996. Effect of precommercial thinning on the production of young fir stands on the Upper North Shore. Canadian Forest Service, Information Report LAU-X-118E, Sainte-Foy, Quebec. 26 pp.nZhang, S. Y. 1995. Effect of growth rate on wood specific gravity and selected mechanical properties in individual species from distinct wood categories. Wood Sci. Technol.29:451-465.nZhang, S. Y. 1997. Variation and correlations of various ring width and ring density features in European Oak: Implications in dendroclimatology. Wood Sci. Technol.31: 63-72.nZhang, S. Y. 1998. Effect of age on the variation, correlations and inheritance of selected wood characteristics in black spruce (Picea mariana). Wood Sci. Technol.32:197-204.nZhang, S. Y., and Z. H. Jiang. 1998. Variability of selected wood characteristics in 40 half-sib families of black spruce (Picea mariana). Wood Sci. Technol.32:71-82.nZhang, S. Y., and E. K. Morgenstern. 1995. Genetic variation and inheritance of wood density in black spruce (Picea mariana) and its relationship with growth: Implications for tree breeding. Wood Sci. Technol.30:63-75.nZhang, S. Y., D. Simpson, and E. K. Morgenstern. 1996. Variation in the relationship of wood density with growth in 40 black spruce (Picea mariana) families grown in New Brunswick. Wood Fiber Sci.28(1):91-99.nZhang, S. Y., Y. Corneau, and G. Chauret. 1998. Impact of precommercial thinning on tree and wood characteristics, and product quality and value in balsam fir. Canadian Forest Service Report No. 39, Forintek Canada Corp., Sainte-Foy, Quebec. 74 pp.nZobel, B. J., and J. P. van Buijtenen. 1989. Wood variation: Its causes and control. Springer-Verlag, Berlin, Germany. 363 pp.n
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